Species of the Moraceae family are of great economic, medicinal and ecological importance in Amazonia. However, there are few studies on their diversity and population dynamics in residual forests. The objective was to determine the composition, structure and ecological importance of Moraceae in a residual forest. The applied method was descriptive and consisted of establishing 16 plots of 20 m × 50 m (0.10 ha), in a residual forest of the Alexánder von Humboldt substation of the National Institute of Agrarian Innovation-INIA, Pucallpa, department of Ucayali, where individuals of arboreal or hemi-epiphytic habit, with DBH ≥ 2.50 cm, were evaluated. The floristic composition was represented by 33 species, distributed in 12 genera; five species not recorded for Ucayali were found. Structurally, the family was represented by 138 individuals/ha with a horizontal distribution similar to an irregular inverted “J”. However, there were different horizontal structures among species. It was determined that 85% of the species were in diameter class I (2.50 to 9.99 cm), being the most abundant Pseudolmedia laevis (Ruiz & Pav.) J.F. Macbr. (41.88 individuals/ha); and the most dominant were Brosimum utile (Kunth) Oken (1.71 m²∕ha) and Brosimum alicastrum subsp. bolivarense (Pittier) C.C.Berg (0.90 m²/ha). Likewise, P. laevis and B. utile were the most ecologically important. The information from the present research will allow the establishment of a baseline, which can be used to propose the management of Moraceae in residual forests in the same study area.

1. Vásquez R, Rojas R, Monteagudo A, et al. Catalog of Peruvian trees. Revista Qeuña 2018; 9(1): 1–167.

2. MINAM (Ministerio del Ambiente, Perú). Plan/Estrategia: Estrategia Nacional de Diversidad Biológica al 2021 (Plan de Acción 2014–2018) (Spanish) [Plan/Strategy: National Biodiversity Strategy to 2021 (Action Plan 2014–2018)]. Lima, Peru: Ministerio del Ambiente; 2014. p. 114.

3. Honorio E, Reynel C. Vacíos en la colección de la flora de los bosques húmedos del Perú (Spanish) [Gaps in the collection of the flora of Peruvian rainforests]. Lima: Universidad Nacional Agraria la Molina, Herbario de la Facultad de Ciencias Forestales; 2003. p. 87.

4. Flores Y. Árboles nativos de la region Ucayali, Perú (Spanish) [Native trees of the Ucayali region, Peru]. 1st ed. Pucallpa, Peru: Instituto Nacional de Innovación Agraria; 2018. p. 354.

5. Mostacedo B; Balcazar J, Montero JC. Tipos de bosque, diversidad y composición florística en la Amazonia sudoeste de Bolivia (Spanish) [Forest types, diversity and floristic composition in the southwestern Amazon of Bolivia]. Ecología en Bolivia 2006; 41(2): 99–116.

6. Calvi SP. Diversidad y distribución de la familia Moraceae en los bosques de la Región Madidi, La Paz, Bolivia (Spanish) [Diversity and distribution of the Moraceae family in the forests of the Madidi Region, La Paz, Bolivia] [PhD thesis]. La Paz: Universidad Mayor de San Andrés; 2013. p. 67.

7. García C, Marín H, Moriones D, et al. Structure, composition and diversity of the natural forests of Smurfit kappa cardboard of Colombia: Popayán and cajibío. Biotecnología en el Sector Agropecuario y Agroindustrial 2014; 12(1): 10–19.

8. Neill D, Killeen T. Curso de dendrología tropical en la Amazonía Boliviana, Valle de Sacta (Spanish) [Tropical dendrology course in the Bolivian Amazon, Sacta Valley]. La Paz: Herbario Nacional de Bolivia; 1991. p. 60.

9. Nebel G, Dragsted J, Vanclay JK. Estructura y composición florística del bosque de la llanura aluvial inundable de la Amazonía Peruana: II El sotobosque de la resting (Spanish) [Structure and floristic composition of the alluvial floodplain forest of the Peruvian Amazon: II The understory of the resting forest]. Folia Amazónica 2000; 10(1–2): 151–181. doi: 10.24841/fa.v10i1-2.246.

10. Cardona V, Fuentes A; Cayola L. Las moráceas de la región de Madidi, Bolivia (Spanish) [The moraceae of the Madidi region, Bolivia]. Ecología en Bolivia 2005; 40(3): 212–264.

11. Araujo-Murakami A, Bascopé F, Cardona V, et al. Composición florística y estructura del bosque amazónico preandino en el sector del Arroyo Negro, Parque Nacional Madidi, Bolivia (Spanish) [Floristic composition and structure of the pre-Andean Amazonian forest in the Arroyo Negro sector, Madidi National Park, Bolivia]. Ecología en Bolivia 2005; 40(3): 281–303.

12. Marcelo-Peña JL, Reynel C. Diversity patterns and floristic composition of permanent evaluative plots in the peruvian central forest. Rodriguésia 2014; 65(1): 35–47. doi: 10.1590/S2175-78602014000100003.

13. Mena-Mosquera VE, Andrade HJ, TorresTorre JJ. Floristic composition, structure and diversity of the tropical pluvial forest of the sub-basin of the Munguidó River, Quibdó, Chocó, Colombia. Entramado 2020; 16(1): 204–215. doi: 10.18041/1900-3803/entramado.1.6109.

14. Torres Navarrete B, Garcia WO. Composición florística de la familia Moraceae, como fuente de carbon aéreo en la gradiente altitudinal de un bosque siempreverde, piemontano de la Amazonia Ecuatoriana, año 2018 (Spanish) [Floristic composition of the Moraceae family, as a source of aerial carbon in the altitudinal gradient of an evergreen, piedmont forest of the Ecuadorian Amazon, year 2018] [Master’s thesis]. Quevedo, Ecuador: Universidad Tecnica Estatal de Quevedo; 2019. p. 95.

15. Brako L, Zarucchi JL. Catálogo de las Angiospermas y Gimnospermas del Perú (Spanish) [Catalog of the Angiosperms and Gymnosperms of Peru]. Missouri: Missouri Botanical Garden; 1993. p. 1–1286.

16. Ulloa Ulloa C, Zarucchi JL, León B. Diez años de Adiciones a la Flora del Perú: 1993–2003 (Spanish) [Ten Years of Additions to the Flora of Peru: 1993–2003]. Arnaldoa 2004; Sp. Ed.: 1–242.

17. Ulloa Ulloa C, Acevedo-Rodríguez P, Beck S, et al. 2017. An integrated assessment of the Vascular plant species of the Americas. Science 2017; 358(6370): 1614–1617. doi: 10.1126/science.aao0398.

18. Berg C, Homeier J. Three new species of South American Moraceae. Blumea-biodiversity, evolution and biogeography of plants 2010; 55(2): 196–200. doi: 10.3767/000651910X527707.

19. Mitidieri N, Cardoso L, Damián A, et al. A new species and a new record of Ficus sect. Pharmacosycea (Moraceae) from Peru. Systematic Botany 2020; 45(1): 91–95.doi: 10.1600/0363636464420X15801369352342.

20. Reynel C, Pennington TD, Pennington RT, et al. Árboles del Perú (Spanish) [Trees of Peru]. Lima: Jesús Bellido M.; 2016. p. 1047.

21. SERFOR (Servicio Nacional Forestal y de Fauna Silvestre, Perú). Anuario forestal y de fauna silvestre 2019 (Spanish) [Forestry and wildlife yearbook 2019]. Lima: SERFOR; 2000. p. 132.

22. FAO (Organización de las Naciones Unidas para la Alimentación y la Agricultura). La Industria de la Madera en el Perú: Identificación de las barreras y oportunidades para el comercio interno de productos responsables de madera, provenientes de fuentes sostenibles y legales, en las MIPYMEs del Perú (Spanish) [The timber industry in Peru: Identification of barriers and opportunities for domestic trade of responsible wood products from sustainable and legal sources in Peruvian MSMEs]. Lima: FAO; 2018. p. 178.

23. Mejía K, Rengifo E. Plantas medicinales de uso popular en la Amazonía peruana (Spanish) [Medicinal plants for popular use in the Peruvian Amazon]. 2nd ed. Lima, Peru: IIAP (Instituto de Investigaciones de la Amazonía Peruana); 2000. p. 286.

24. Mass W, Campera M. Árboles medicinales: Conocimientos y usos en la cuenca baja del río Marañón, zona de amortiguamiento de la Reserva Nacional Pacaya Samiria (Spanish) [Medicinal trees: Knowledge and uses in the lower basin of the Marañón River, buffer zone of the Pacaya Samiria National Reserve]. Iquitos: MINAM; 2011. p. 81.

25. Spichiger R, Méroz J, Loizeau P, et al. Contribución a la flora en la Amazonía peruana: Los árboles del Arboretum de Jenaro Herrera. Volumen II: Linaceae a Palmae (Spanish) [Contribution to the flora in the Peruvian Amazon: The trees of the Jenaro Herrera Arboretum. Volume II: Linaceae to Palmae]. Geneva, Switzerland: IIAP (Instituto de Investigaciones de la Amazonía Peruana); 1990. p. 359.

26. Shanahan M, Samson SO, Estephen SG, et al. Fig-eating by vertebrate frugivores: A global review. Biological Reviews 2001; 76(4): 529–572. doi: 10.1017/S146479310100576010.

27. Kanashiro LJ. Etología de Forrajeo de Ateles belzebuth chamek (Atelidae: Atelinae) en el Parque Nacional del Manu durante la temporada seca 2005 (Spanish) [Foraging ethology of Ateles belzebuth chamek (Atelidae: Atelinae) in Manu National Park during the 2005 dry season.] [PhD thesis]. Lima: Universidad Nacional Agraria La Molina; 2009.

28. Alegria DO. Influencia de la disponibilidad de frutos (familia Moraceae) en las dinámicas de fisión-fusión de Ateles chamek (Humboldt, 1812) en el Parque Nacional de Manu (Spanish) [Influence of fruit availability (Moraceae family) on fission-fusion dynamics of Ateles chamek (Humboldt, 1812) in Manu National Park] [PhD thesis]. Lima: Universidad Nacional Agraria La Molina; 2019. p. 75.

29. ter Steege H, Pitman N, Sabatier D, et al. 2013. Hyperdominance in the Amazonian tree flora. Science 2013; 342(6156): 245–337.

30. a)doi: 10.1126/science.1243092.

31. Licona JC, Peña M, Mostacedo B. Composición florística, estructura y dinámica de un bosque amazónico aprovechado a diferentes intensidades en Pando, Bolivia (Spanish) [Floristic composition, structure and dynamics of an Amazonian forest harvested at different intensities in Pando, Bolivia]. Santa Cruz, Bolivia: Instituto Boliviano de Investigación Forestal; 2007. p. 60.

32. FAO (Naciones Unidas para la Alimentación y la Agricultura, Italia); SERFOR (Servicio Nacional Forestal y de Fauna Silvestre, Perú). Nuestros bosques en números: Primer reporte del Inventario Nacional Forestal y de Fauna Silvestre (Spanish) [Our forests in numbers: First report of the National Forest and Wildlife Inventory] [Internet]. Lima: FAO, SERFOR; 2017. Available from: https://sinia.minam.gob.pe/documentos/nuestros-bosques-numeros.

33. Vidaurre HE. Balance of silvicultural experience with cedrelinga catenaeformis Ducke in the Pucallpa region in the Peruvian Amazone [Master’s thesis]. Turrialba, Costa Rica: CATIE; 1994. p. 165.

34. Angulo W, Fasabi H. Fenología de 10 especies forestales para determinar la influencia del cambio climático por efecto del calentamiento global: cinco años de estudio (2012–2016) (Spanish) [Phenology of 10 forest species to determine the influence of climate change due to the effect of global warming: Five years of study (2012–2016)]. Pucallpa: INIA; 2016. p. 31.

35. Bridson D, Forman L. The herbarium handbook. Kew: Royal Botanic Gardens; 1992. p. 93.

36. Lamprecht H. Silvicultura en los trópicos. Los ecosistemas forestales en los bosques tropicales y sus especies arbóreas. Posibilidades para un aprovechamiento sostenido (Spanish) [Silviculture in the tropics. Forest ecosystems in tropical forests and their tree species. Possibilities for a sustainable use]. Carrillo A (translator). Eschborn, Federal Republic of Germany: GTZ; 1990. p. 335.

37. Finegan B. El potencial de manejo de los bosques húmedos secundarios neotropicales de las tierras bajas (Spanish) [The management potential of lowland neotropical secondary moist forests]. Luján R (translator). Turrialba: CATIE; 1992. p. 37.

38. Mostacedo B, Fredericksen T. Manual de métodos básicos de muestreo y análisis en ecología vegetal (Spanish) [Manual of basic methods of sampling and analysis in plant ecology]. Santa Cruz de la Sierra: BOLFOR; 2000. p. 87.

39. Colwell RK, Chang XM, Jing C. Interpolando, extrapolando y comparando las curvas de acumulación de especies basadas en su incidencia (Spanish) [Interpolating, extrapolating and comparing species accumulation curves based on their occurrence]. Ecology 2005; 85(10): 2717–2727.

40. Magurran EA. Measuring biological diversity. Oxford: Blackwell Publishing; 2004. p. 256.

41. Colwell RK. Estimates 9.1.0 user’s guide: Statistical estimation of species richness and shared species from samples. Connectitut: University of Connecticut; 2013.

42. Louman B, Quirós D, Nilsson M. Silvicultura de bosques latifoliados húmedos con énfasis en América Central (Spanish) [Silviculture of moist broadleaf forests with emphasis on Central America]. Turrialba: CaTIE; 2001. p. 265.

43. Curtis JT, McIntosh RP. An upland forest continuum in the prairieforest border region of Wisconsin. Ecological Society of America 1951; 32(3): 476–496. doi: 10.2307/1931725.

44. Orozco L, Brumér L. Inventarios forestales para bosques latifoliados en América Central (Spanish) [Forest inventories for broadleaved forests in Central America]. Turrialba: CATIE; 2002. p. 264.